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U l t i ma t e l y the p l e u r o p e r i t o n e a l a n d pleuropericardial folds separate the pericardioperitoneal canals from the peritonea a n d p e r i c a r d i a l c a v i t i e s, r e s p e c t i v e l y, a n d the r e ma i n i n g s p ap re s fi oirv e the c im t m p l e u r a l c a v i t i e s e eC h a p t e r 11 the me s o d e r m, w h i c h c o v e r s the o u t s i d e o f the (s). E xp a n s i o n o f the l u n g b u d s i n t o the p e r i c a r d i o p e r i t o n e a l c a n a l s. At 6 t h i s s t a g e the c a n a l s a r e i n c o mmu n i c a t i o n w i t h the p e r i t o n e a l a n d p e r i c a r d i a l c a v i t i e sA. T buds showing the pleuropericardial folds that will divide the thoracic portion of the body cavity into the pleural and pericardial cavities. D u r i n g f u r the r d e v e l o p me n t, s e c o n d a r y b r o n c h i d i v i d e r e p e a t e d l y i n a d i c h o t o mo u s f a s h i o n, f o r mi n g t e n t i a r y (s e g m e n t b rl o n c h i i n the r i g h t l u n g a n d e i g h t i n the r a) l e f t, c r e a t i n g tb r o n c h o p u l m o n a r y s e g m eo f t sh e a d u l t l u n g. B y the e n d o f he n t the s i xt h mo n t h, a p p r o xi ma t e l y 1 7 g e n e r a t i o n s o f s u b d i v i s i o n s h a v e f o r me d. B e f o r e the b r o n c h i a l t r e e r e a c h e s i t s f i n a l s h a p e, a n w e v ei r,i o n a l s i x d i v i s i o n s ho add t f o r m d u r i n g p o s t n a t a l lB f e n c h i n g i s r e g u l a t e d b y e p i the l i a l - me s e n c h y ma l ira. S i g n a l s f o r b r a n c h i n g, w h i c h e mi t f r o m the me s o d e r m, i n v o l v e me mb e r s o f the f i b r o b l a s t g r o w t h f a c t o r f a mi l y. 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T h e c e p h a l i c e n d o f the n e u r a l t u b e s h o w s t h r e e d i l p t iio na r yt h er a i n ar m s, b v e s i c l e s:a) t h eP r o s e n c e p h a l o n r f o r e b r a i n(;b) t h eM e s e n c e p h a l o n r (, o, o m i d b r a i n; n d c) t h eR h o m b e n c e p h a l,o n r h i n d b r a i n i g. S i mu l t a n e o u s l y i t f o r ms t w o f l e xu r the sec(e r v i c a l f l e x u r e the j u n c t i o n o f the a) h: at h i n d b r a i n a n d the s p i n a l c o r db) tnh ec e p h a l i c f l e x u rn the mi d b r a i n r e g i o n a d (ie (s e eF i g. W h e n the e mb r y o i s 5 w e e k s o l d, the p r o s e n c e p h a l o n c o n s i s t s o f t w e p a r t s: (a) h o The l e n c e p h a l o no r me d b y a mi d p o r t i o n a n d t w o l a t e r a l o u t p o c k e t i n g s, the, f p r i m i t i v e c e r e b r a l h e m i s p h earn d b) t h eD i e n c e p h a l o n h a r a c t e r i ze d b y, es (, c o u t g r o w t h o f the o p t i c v e s i c l e s i (gs. T h e r h o mb e n c e p h a l o n a l s o c o n s i s t s o f t w o pta reM:e (t e n c e p h a l o n h i c h a) h t s, w l a t e r f o r ms the n s a n dc e r e b e l l u ma n d b) t h em y e l e n c e p h a l o n. T h e c a v i t y o f the r h o mb e n c e p h afl o u r its t v e n t r i c l the a t o f n h h, h the d i e n c e p h a l o n i s tth ie d v e n t r i c l e n d t h o s e o f the c e r e b r a l h e mi s p h e r e s a r e hr, a the l a t e r a l v e n t r i c l (Fsg. T h i s l u me n b e c o me s v e r y n a r r o w a n d i s the n k n o w n a s the a q u e d u c t o f S y l v i u s. C o l l e c t i v e l y the y c o n s t i t u t e the n e u r o e p i the l i a l l a y en e u r o e p i the l i u m. T h e y f o r m the n t l e l a y e r zo n e a r o u n d the n e u r o e p i the l i a l l a y e r (s e e) Ma, a F i g. D o r s a l v i e w o f a l a t e p r e s o mi t e e mb r y o a t a p p r o xi ma t e l y 1 8 1 d a y s. T h e a mn i o n h a s b e e n r e mo v e d, a n d the n e u r a l p l a t e i s c lB a r l y v i s i b l. T r a n s v e r s e s e c t i o n s t h r o u g h s u c c e s s i v e l y o l d e r e mb r y o s 2 s h o w i n g f o r ma t i o n o f the n e u r a l g r o o v e, n e u r a l t u b e, a n d n e u r a l c r e s t. C e l l s o f the n e u r a l c r e s t mi g r a t e f r o m the e d g e s o f the n e u r a l f o l d s a n d d e v e l o p i n t o s p i n a l a n d c r a n i a l s e n s o r y g a n g lCa D. D o r s a l v i e w o f a h u ma n e mb r y o a t a p p r o xi ma t e l y d a y 2 2. S e v e n 3 d i s t i n c t s o mi t e s a r e v i s i b l e o n e a c h s i d e o f the nB. T h e n e r v o u s s y s t e m i s i n c o n n e c t i o n w i t h the a mn i o t i c c a v i t y t h r o u g h the c r a n i a l a n d c a u d a l n e u r o p o r e s. D r a w i n g o f a s a g i t t a l s e c t i o n t h r o u g h the b r a i n a t a p p r o xi ma t e l y 2 7 4 d a y s o f h u ma n d e v e l o p me n t. T h r e e b r a i n v e s i c l e s r e p r e s e n t t(h e, f o r e b r a i n F) mi d b r a i n M), a n d h i n d b r a(i H). D r a w i n g o f a s a g i t t a l s e c t i o n t h r o u g h the b r a i n a p p r o xi ma t e l y 3 2 5 d a y s o f h u ma n d e v e l o p me n t. T h e t h r e e o r i g i n a l b r a i n v e s i c l e s h a v e s e g r e g a t e d i n t o the t e l e n c e p h a l o n, d i e n c e p h a l o n, me s e n c e p h a l o n, me t e n c e p h a l o n, a n d my e l e n c e p h a l o n. S e c t i o n o f the w a l l o f 6 neuroepithelial cells, which form a the full width of the wall. Note the S c a n n i n g e l e c t r o n mi c r o g r a p h o f a s i mi l a r t o t h a t Ai. S e c t i o n o f the n e u r a l t u b e a t a s l i g h t l y mo r e a d v a n c e d s t a g e t h a n 7 i n F i g u r e 1 7. O n the p e r i p h e r y, i mme d i a t e l y a d j a c e n t t o the e xt e r n a l l i mi t i n g me mb r a n e, n e u r o b l a s t s f o r m. T h e s e c e l l s, w h i c h a r e p r o d u c e d b y the n e u r o e p i the l i a l c e l l s i n e v e r - i n c r e a s i n g n u mb e r s, w i l l f o r m the ma n t l e l a y e r. As a r e s u l t o f my e l i n a t i o n o f n e r v e f i b e r s, t h i s l a y e r t a k e s o n a w h i t e a p p e a r a n c e a n d the r e f o r e ws ic a l l e d the i h the m a t t e r o f the s p i n a l c(F r g. B a s a l, A l a r, R o o f, a n d F l o o r P l a t e s As a r e s u l t o f c o n t i n u o u s a d d i t i o n o f n e u r o b l a s t s t o the ma n t l e l a y e r, e a c h s i d e o f the neural tube shows a ventral and a dorsal thickening. The ventral thickenings, the B a s a l p l a t e s h i c h c o n t a i n v e n t r a l mo t o r h o r n c e l l s, f o r m the mo t o r a r e a s o f, w the s p i n a l c o r d; the d o r s a l t h i c k e n i n gls,r t p ea t e sf o r m t h s e n s o r y a r e a s A a hl, e (F i g. T h e d o r s a l a n d v e n t r a l mi d l i n e p o r t i o n s o f the n e u r a l t u b e, k n o w n a s t h er o o f a n df l o o r p l a t e s e s p e c t i v e l y, d o n o t c o n t a i n n e u r o b l a s t s; the y s e r v e, r p r i ma r i l y a s p a t h w a y s f o r n e r v e f i b e r s c r o s s i n g f r o m o n e s i d e t o the o the r. In a d d i t i o n t o the v e n t r a l mo t o r h o r n a n d the d o r s a l s e n s o r y h o r n, a g r o u p o f n e u r o n s a c c u mu l a t e s b e t w e e n the t w o a r e a s a n d f o r mst e r mma l il a t e h o r n in a s ed (F i g.

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